Traditionally I thought it was QB, then S (which are kind of the QBs of the secondary)...
BUT, with the limited practice times more and more since 2010, the schemes on both sides have gotten extremely more simpler... and therefore easier on the QBs and S mentally. But still, Safety needs to know the entire defense, maybe not DL, but certainly the rest.
TE with needing to know all the WR and OL stuff is tough too.
That would be an interesting debate for some pros to argue.
Also, I like Jason Kelce, always downplays the Center position compared to the rest of the OL. Saying maybe the snapping is tough for some, but after the snap, Center is easily the easiest OL position.
Most NFL players are always hyping their own positions as one of the toughest, and Kelce is just the opposite, and like, heck no, it's you guys.
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